Some species such as Celtis Africana, are experiencing relative recruitment bottlenecks, because there are usually fewer recruits [i.e. individuals <20 cm diameter at breast height, (dbh)] than canopy individuals. The species with low recruitment tent to have higher concentrations of leaf nitrogen, suggesting that they are shade-intolerant species. Similarly the canopy species which are recruiting adequately have low level of leaf nitrogen.
Reference:
Midgley, JJ, Everard, DA and Vanwyk, G. 1995. Relative lack of regeneration of shade-intolerant canopy species in some South African forests. South African Journal of Science, vol. 91(1), pp 7-8
Midgley, J., Everard, D., & Vanwyk, G. (1995). Relative lack of regeneration of shade-intolerant canopy species in some South African forests. http://hdl.handle.net/10204/979
Midgley, JJ, DA Everard, and G Vanwyk "Relative lack of regeneration of shade-intolerant canopy species in some South African forests." (1995) http://hdl.handle.net/10204/979
Midgley J, Everard D, Vanwyk G. Relative lack of regeneration of shade-intolerant canopy species in some South African forests. 1995; http://hdl.handle.net/10204/979.